Theropoda

    The theropods are somewhat bland on the surface, because they all followed a body plan devoted to bipedal hunting on the ground, but they had a healthy variety of styles and variations on their main theme.  It is fairly well settled that birds are closely related to theropods, although the placement and groups closest are not clear.  I used to follow a theory that claims flight was achieved by animals just before the first true dromaeosaurids, making a number of classic (historically considered non-avian and not having ancestors that could fly) theropods secondarily flightless, meaning they had ancestors that could fly, but they themselves had lost that ability.  This is between two other theories, known colloquially as BADD (Birds Are Dinosaur Descendants, with the emphasis here that no classic theropod was secondarily flightless) and BCF (Birds Came First, with the premise that all classic theropods were secondarily flightless).  Since I've gotten my copy of The Dinosauria II, I've finally come around to BADD.  
    The theropods were all bipedal, as far as is known, and the majority of the classic theropods were predators, with some exceptions: the oviraptorosaurians, troodontids, and ornithomimosauroids may have been herbivorous or omnivorous.  They all likely could move reasonably fast; the large theropods had their speed helped by their long legs.  By taking large strides quickly, they could move fast without running if they chose.  It is likely that at least all theropods that were members of Coelurosauria either were covered with a downy "protofeather" coating or had lost that characteristic. 
    Some important anatomical characters linking all theropods ancestrally include an extra hole in the skull's maxilla, a reduction or loss of both the fourth and fifth digit of the hand, hollow long bones, reduction of the fifth digit of the foot to almost nothing, and presacrals bearing holes (pleurocoels) to hollow centra.
    In the past, a group called the teratosaurids was considered to be the earliest "carnosaurs," and sometimes was aligned with the prosauropods.  This group was eventually discovered to be based on "thecodont" remains, sometimes mixed with prosauropods remains for good measure, but not before working its way into the popular literature.  A fair number of older (pre~1985) books on dinosaurs include (very hypothetical) restorations of "teratosaurids". 
    Although it was recently thought that a sauropod, Eobrontosaurus, had "belly ribs," it appears that only theropods and prosauropods had these, which would have protected the belly area.  In addition, many theropods have been restored with their palms facing down or back, which would have been impossible due to lack of the proper forearm rotation for most (if not all) of them.  Instead, theropod hands would have faced each other.
    An interesting possibility suggests that the adults of small theropods may have had to compete with the juveniles of large theropods, particularly if the parents had left the young to fend for themselves at an early age.  This in effect would have made these juveniles almost a separate species.  Theropods may have also done communal nesting, as suggested by remains found in Portugal.   
    An unusual small tooth from the Late Campanian of Mexico has a peculiar groove that could indicate the existence of a small, venomous theropod.

<--Theropoda
      |--?Guaibasaurus
  
   `-->Neotheropoda

Theropoda:

Taxon or Taxa: Time\Place: Comments:
Guaibasaurus candelariai Bonaparte, Ferigolo, and Ribeiro, 1999 earliest Carnian (LTr) of Brazil This new basal dinosaur has been considered to be somewhere around Staurikosaurus and the rest of Herrerasauridae, or even basal to the Theropoda\ Sauropodomorpha split.  Alternately, it may instead be basal to Neotheropoda, as shown here.  Remains of two individuals are known, with the type including several dorsals, two sacrals, 10 caudals, ribs and chevrons, most of the hip and shoulder girdles, and partial hindlimbs.  The acetabulum shows little opening, but the fifth toe shows reduction.

Theropoda i.s.:  Don't weep for most of these folks.  Aside from a couple of animals that appear to belong around this level, these creatures earned their status, being based on little material, often poor in quality.  More than a few tooth taxa appear in this following table, providing mute testimony to the fact that dinosaurs named from isolated teeth more often than not quickly enter into a usually richly-deserved obscurity.  Occasionally, one will rise from the depths like Troodon on the basis of uniqueness, but this is rare (in the old days of dinosaur paleo, a dubious tooth name like Trachodon could became host to a taxonomical monstrosity of names and material, very little of which could ever be shown to pertain to the host taxon).

Taxon or Taxa: Time\Place: Comments:
"Allosaurus" sibiricus (N.D.) Riabinin, 1914 Berriasian-Hauterivian (EK) of Siberian Russia This obscure species is based on a undiagnostic portion of metatarsal.
Chienkosaurus ceratosauroides (N.D.) Yang, 1942 ?LJ of China Based on four teeth, three of which turned out to be crocodilian, and the other of which is usually thrown in with Szechuanosaurus campi, Chienkosaurus is probably not assignable to it, simply because the tooth is pretty much the same as those from Yangchuanosaurus and Sinraptor, too.  It was originally thought to be close to Ceratosaurus, hence the species name.
?Chingkankousaurus fragilis (N.D.) Yang, 1958 early-mid Maastrichtian (LK) of China Based on a supposedly tyrannosaurid-type scapula that could just as well be a partial dorsal rib, Chingkankousaurus cannot be classified any farther than Theropoda, if you're feeling charitable.
"Creosaurus" potens (N.D.) Lull, 1911 late Aptian-early Albian (EK) of Maryland "Creosaurus" potens is based on an unusual caudal vert.
Dandakosaurus indicus (N.D.) Yadagiri, 1990 Pliensbachian-Toarcian (EJ) of India So far, this is a very mysterious animal.  Little has yet been reported.  It may be a ceratosaurian, or basal tetanuran.
"Elaphrosaurus" iguidiensis (N.D.) Lapparent, 1960 late Aptian-Albian (EK) of Algeria, Libya, and Niger This species cannot be assigned with certainty to Elaphrosaurus.  I wouldn't be surprised if some of the material (including a pubic boot with a distinctive morphology) assigned here belongs to abelisauroids, spinosaurids, or carcharodontosaurids.
Embasaurus minax (N.D.) Riabinin, 1931 Berrisian-Hauterivian (EK) of Kazakhstan Both a geographic curiosity and poorly known, the vertebrae this form is based on could belong to a basal tetanuran.
?Inosaurus tedreftensis (N.D.) Lapparent, 1960 Albian (EK) of Niger There's no proof that the seemingly arbitrarily-assigned remains named Inosaurus actually belong together.  It is based on twenty-five small but heavily built vertebrae and part of a tibia, recovered from three different localities.  Inosaurus may well be chimerical and from the description it sounds as if the verts could just as well belong to a juvenile sauropod.
"Kagasaurus" (N.N.) Hisa, 1989 EK of Japan "Kagasaurus", based on two teeth, may be the same as Fukuiraptor.
"Katsuyamasaurus" (N.N.) Lambert, 1990 EK of Japan The name is interesting.  This animal, based on an ulna and a caudal, may be a Sinraptor-Allosaurus type theropod, possibly the same as Fukuiraptor.
"Liassaurus huenei" (N.N.) Welles, Powell, and Pickering vide Pickering, 1995 (?Sarcosaurus) Sinemurian (EJ) of England This theropod is based on a collection of hindlimb and pelvic elements.  These bones are from the same time and place as Sarcosaurus, and very similar to boot, so may belong to it. 
"Megalosaurus": "M." chubutensis (N.D.) Corro, 1974 LK of Argentina This is an indeterminate tooth species that can't be assigned to the European MJ Megalosaurus, or any other theropod for that matter.
"M." dunkeri (N.D.) Dames, 1884 Berriasian-?Barremian (EK) of Germany, ?England, and ?Spain Much more famous as its alter-ego Altispinax, "M." dunkeri is a tooth species from Germany that cannot be shown to have any connection to the three tall vertebrae assigned to it by Huene that formed the basis of the name Altispinax and now are known as Becklespinax.  It used to be illustrated as having a sort of hump near the shoulders.  It may be related to Prodeinodon and/or a maniraptoran.
"M." hungaricus (N.D.) Nopcsa, 1901 early-mid Maastrichtian (LK) of Romania This is just another tooth species.
"M." inexpectatus (N.D.) Corro, 1966 ?Coniacian-?Maastrichtian (LK) of Argentina The teeth this species is based on could belong to an abelisauroid or carcharodontosaurid by time and place.
"M." insignis (N.D.) Eudes-Deslongchamps and Lennier, 1870 Oxfordian-Valanginian (LJ-EK) of France, Portugal, and ?England This species is based on teeth and postcrania, and additional remains are referred to it, but on what basis I don't know.
"M." lonzeensis (N.D.) Dollo, 1883 Santonian (LK) of Belgium This isolated pedal ungual is not referable to Megalosaurus.
"M." lydekkeri (N.D.) Huene, 1926 Sinemurian (EJ) of England This is (surprise) another doubtful tooth species.
"M." pannoniensis (N.D.) Seeley, 1881 Campanian-Maastrichtian (LK) of Austria, France, and Portugal This is just another doubtful tooth species.
"M." pombali (N.D.) Lapparent and Zybszewski, 1957 Oxfordian-Kimmeridgian (LJ) of Portugal For a change of pace, this indeterminate species is based on vertebrae.
"M." terquemi (N.D.) Huene, 1926 Hettangian (EJ) of France We (finally!) reach the last of this batch of dubious Megalosaurus species, ending with another species based on teeth.
"Mifunesaurus" (N.N.) Hisa, 1989 Cenomanian (LK) of Japan This is another undescribed theropod.
"Morosaurus" marchei (N.D.) Sauvage, 1897-98 Aptian-Cenomanian (EK-LK) of Portugal This is a dubious theropod based on a distal caudal.
"Ngexisaurus dapukaensis" (N.N.) Zhao, 1983 MJ of China Very little, beyond the interesting name, is known of this dinosaur, which may be a coelurosaurian or, according to the faunal list in The Dinosauria II, a ceratosaurian.
?"Ornithocheirus" hilesnis (N.D.) Koken, 1885 EK of Germany Based on a toe bone, this was originally described as a species of the pterosaur Ornithocheirus, but it's more likely to have been a theropod.  Unfortunately, the bone is now lost.
Ornithodesmus cluniculus (N.D.) Seeley, 1887 Barremian (EK) of England Long confused with birds and pterosaurs, the type sacrum of this animal has been assigned to a troodontid or dromaeosaurid in the past, but new restudy suggest it might not even be tetanuran.
Orthogoniosaurus matleyi (N.D.) Das Gupta, 1931 mid-late Maastrichtian (LK) of India This theropod is based on an odd tooth.
Phaedrolosaurus ilikensis (N.D.) Dong, 1973 ?Valanginian-Albian (EK) of China Little is known about this theropod, which some have speculated is close to the ancestry of Dromaeosauridae.  It is based on a somewhat dromaeosaurid-like but rather uninformative tooth.  Referred non-dromaeosaurid but maniraptoran hindlimb material has been shown to be diagnostic and has been renamed Xinjiangovenator.
"Poekilopleuron" schmidti (N.D.) Kiprijanov, 1883 Cenomanian (LK) of western Russia The remains named "Poekilopleuron" schmidti could belong to any large LK theropod (provided it had cause to be passing through the Cenomanian of western Russia).
Prodeinodon mongoliensis (N.D.) Osborn, 1924 Barremian-early Aptian (EK) of Mongolia This tooth taxon is based on some choppers with a resemblance to those named Aublysodon, suggesting possibly a young tyrannosaurid.  It may, though, be a maniraptoran.
"Prodeinodon" kwangshiensis (N.D.) Hou, Yeh, and Zhao, 1975 Barremian-early Aptian (EK) of China This is a tooth taxon that may be close to the similarly poorly-known Wakinosaurus.
Protoavis texensis Chaterjee, 1991 late Carnian (LTr) of Texas Protoavis is a...thing, perhaps even a "fiendish thingie", in the words of George Harrison, and well-known for its charming habit of presenting a different face to everyone who looks at it.  This dinosaur represents a great controversy.  Some have called it an early bird, while others have considered it a chimerical muddle.  Its remains have been accused of coming from a combination of coelophysoids, herrerasaurids, early coelurosaurs, and pterosaurs and various other nondinosaurians.  However, it is now beginning to look as though much of the material goes together (although some was misidentified originally).  My best guess is that this animal will turn out to be closest to the coelophysoids.
New restorations show a long forelimb, but the shoulder, hindlimb, and pelvis seem poorly known.  Although many bones are known, the skeleton is still fragmentary and open to interpretation.  Apparently, part of the hindlimb is coelophysoid, but parts of the rest of it are not, and I'll hold off putting it in one place until a type is decided upon.
Siamosaurus suteethorni (N.D.) Buffetaut and Ingavat, 1986 LJ or EK of Thailand This is a possible spinosaurid based on teeth, although even its dinosaurian status is questionable.
Sinocoelurus fragilis (N.D.) Yang, 1942 Tithonian (LJ) of China This tooth species has unserrated teeth reminiscent of spinosaurids, so it may be a small or juvenile early member.  It also may not be.  Let it be known that I am the one who started this poorly-founded rumor.
Szechuanosaurus campi (N.D.) Yang, 1942 Oxfordian (LJ) of China Szechuanosaurus is a name you may run across that isn't very useful.  It was originally based on some shed teeth.  Later, a headless partial skeleton was found and referred to this taxon ("S." zigongensis).  Unfortunately, there is no conclusive way to link the teeth and skeleton, theropod teeth being what they are.
"Szechuanosaurus" yandonensis (N.D.) Dong, Chang, Li, and Zhou, 1978 Oxfordian (LJ) of China This is a dubious theropod based on teeth.
?"Tanystrophaeus" posthumus (N.D.) Huene, 1908 mid Norian (LTr) of Germany Also known as Tanystrosuchus, this is just an obscure indeterminate ?theropod based on a caudal.
Teinurosaurus sauvagei (N.D.) Nopcsa, 1928 Kimmeridgian (LJ) of France The caudal vert this species is based on has gone through a taxonomic mess.  Nopcsa, by typographical error, almost named it Saurornithoides, which wouldn't have mattered as much if that name wasn't already in use.  Understandably, he quickly printed a clarification, which von Huene missed on his way to naming it Caudocoelus in 1932.  It's dubious by any name, and was destroyed in WWII to boot.
"Tomodon" horrificus (N.D.) Leidy, 1865 (?Dryptosaurus) ?Maastrichtian (LK) of New Jersey This animal, better known as Diplotomodon (Tomodon, it turned out, was preoccupied), is possibly the same as Dryptosaurus.
"Tyrannosaurus lanpingensis" (N.N.) Yeh, 1975 Berriasian-Hauterivian (EK) of China This is an indeterminate tooth species, far too old to be Tyrannosaurus.
Wakinosaurus satoi (N.D.) Okazaki, 1992 Berriasian-?Hauterivian (EK) of Japan Described as a "megalosaurid," Wakinosaurus satoi is based on teeth.
Walgettosuchus woodwardi (N.D.) Huene, 1932 Albian (EK) of Australia This species is based on a poorly preserved caudal vertebra that could belong to any one of several types of theropods
Yaverlandia bitholus Galton, 1971 Barremian (EK) of England An unusual Wealden dinosaur, Yaverlandia is based on a skull roof showing thickening, which led to it becoming famous as possibly the earliest, most generalized pachycephalosaurid. However, upon further review, it isn't a pachycephalosaur, and actually shows theropod characteristics.
"Zanclodon" cambrensis" (?N.D.) Newton, 1899 Rhaetian (LTr) of Wales Based on a partial jaw, this animal was for a long time assigned to Megalosaurus.  However, it appears to be closer to the coelophysoids than to the tetanurans.  It could be indeterminate. 

Neotheropoda: This is getting into the more derived theropods, and starts off with a split between the coelophysoids and the other theropods, followed closely by the splitting-off of Ceratosauria.

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